Hear it pronouncedPeregrine Falcon
| Peregrine Falcon | ||||||||||||||
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 Adult of subspecies pealei or tundrius, Alaska
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| Falco peregrinus Tunstall, 1771 |
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 Global range
Yellow: Breeding summer visitor Green: Breeding resident Blue: Winter visitor Light blue: Passage visitor |
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17-19, see text |
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Falco atriceps Hume |
The Peregrine Falcon (Falco peregrinus), also known simply as the Peregrine,[2] and historically as the "Duck Hawk" in North America,[3] is a cosmopolitan bird of prey in the family Falconidae. It is a large, crow-sized falcon, with a blue-gray back, barred white underparts, and a black head and "moustache". It can reach speeds over 322 km/h (200 mph), making it the fastest animal in the world.[4] As is common with bird-eating raptors, the female is much bigger than the male.[5][6] Experts recognize 17–19 subspecies, which vary in appearance and range; there is disagreement over whether the distinctive Barbary Falcon is a subspecies or a distinct species.
The Peregrine's breeding range includes land regions from the Arctic tundra to the Tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread bird of prey.[7] Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations.
While its diet consists almost exclusively of medium-sized birds, the Peregrine will occasionally hunt small mammals, small reptiles or even insects. It reaches sexual maturity at one year, and mates for life. It nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures.[8] The Peregrine Falcon became an endangered species in many areas due to the use of pesticides, especially DDT. Since the ban on DDT from the beginning of the 1970s onwards, the populations recovered, supported by large scale protection of nesting places and releases to the wild.[9]
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[ Description
The Peregrine Falcon has a body length of 34 to 58 centimetres (13–23 in) and a wingspan of around 80 to 120 centimetres (31–47 in).[5][10] The male and female have similar markings and plumage, but as in many birds of prey the Peregrine Falcon displays marked reverse sexual dimorphism in size, with the female measuring up to 30 percent larger than the male.[11] Males weigh 440–750 g, and the noticeably larger females weigh 910–1500 g; for variation in weight between subspecies, see under that section below.
The back and long, pointed wings of the adult are usually bluish black to slate gray with indistinct darker barring (see "Subspecies" below); the wingtips are black.[10] The underparts are white to rusty and barred with thin clean bands of dark brown or black.[12] The tail, colored like the back but with thin clean bars, is long, narrow and rounded at the end with a black tip and a white band at the very end. The top of the head and a "mustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat.[13] The cere is yellow, as are the feet, and the beak and claws are black.[14] The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck.[4][5][6] The immature bird is much browner with streaked, rather than barred, underparts, and has a pale bluish cere.[5][13]
[ Taxonomy and systematics
This species was first described by Marmaduke Tunstall in his 1771 Ornithologia Britannica under its current binomial name.[15] The scientific name Falco peregrinus, means "wandering falcon" in Latin.[16] Indeed, the species' common name refers to its wide-ranging flights in most European languages.[17] The Latin term for falcon, falco, is related to falx, the Latin word meaning sickle, in reference to the silhouette of the falcon's long, pointed wings in flight.[4]
The Peregrine Falcon belongs to a genus whose lineage includes the hierofalcons[18] and the Prairie Falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Early Pliocene, about 8–5 million years ago (mya). As the Peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear; the issue is complicated by widespread hybridization confounding mtDNA sequence analyses; for example a genetic lineage of the Saker Falcon (F. cherrug) is known[19] which originated from a male Saker producing fertile young with a female Peregrine ancestor some 100,000 years ago.[ref]
Today, Peregrines are regularly hybridized in captivity with other species such as the Lanner Falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the Peregrine's hunting skill with the Lanner's hardiness, or the Gyrfalcon to produce large, strikingly colored birds for the use of falconers. As can be seen, the Peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).[20]
[ Subspecies
Numerous subspecies of the Peregrine have been described, with 19 accepted by the Handbook of the Birds of the World.[21][5][6]
- Falco peregrinus peregrinus, the nominate subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south.[22] It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580–750 g, while females weigh 925–1,300 g.[6] It includes brevirostris, germanicus, rhenanus, and riphaeus.
- Falco peregrinus calidus, described by John Latham in 1790, was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia, from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory, and travels south in winter as far as sub-Saharan Africa. It is paler than peregrinus, especially on the crown. Males weigh 588–740 g, while females weigh 925–1,333 g.[6]
- Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti and pleskei, and harterti seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on coast there), and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to peregrinus, but the young are even darker than those of anatum.
- Falco peregrinus macropus, described by Swainson in 1837 is the Australian Peregrine Falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.[12]
- Falco peregrinus submelanogenys described by Mathews in 1912, is the Southwest Australian Peregrine Falcon. It is found in southwest Australia and is non-migratory.
- Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian Peregrine Falcon, Black Shaheen, or Indian Shaheen.[23] It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from Pakistan across India to Sri Lanka and Southeastern China; in Pakistan it is a military symbol of the Pakistan Air Force. It is non-migratory. It is small and dark, with rufous underparts barred with lighter color. In Sri Lanka this species is found to favour the higher hills while the migrant calidus is more often seen along the coast.[24]
- Falco peregrinus anatum, described by Bonaparte in 1838,[22] is known as the American Peregrine Falcon, or "Duck Hawk"; its scientific name means "Duck Peregrine Falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains today. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts seek to restore the population.[22] Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to peregrinus but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500–570 g, while females weigh 900–960 g.[25][5]
- Falco peregrinus cassini, described by Sharpe in 1873, is also known as the Austral Peregrine Falcon. It includes kreyenborgi, the Pallid Falcon[26] a leucistic morph occurring in southernmost South America, which was long believed to be a distinct species.[27] Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and Falkland Islands.[12] It is non-migratory. It is similar to nominate, but slightly smaller with a black ear region. The variation kreyenborgi is medium grey above, has little barring below, and has a head pattern like the Saker Falcon, but the ear region is white.[27]
- Falco peregrinus pealei, described by Ridgway in 1873, is also known as Peale's Falcon, and includes rudolfi.[28] It is found in the Pacific Northwest of North America, northwards from the Puget Sound along the British Columbia coast (including the Queen Charlotte Islands), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia.[28] It is possibly found on the Kuril Islands and the coasts of Kamchatka as well. It is non-migratory. It is the largest subspecies, and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide.[29] Juveniles occasionally have pale crowns.
- Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys It is found in the Arctic tundra of North America to Greenland. It migrates to wintering grounds in Central and South America.[29] Most vagrants that reach western Europe belong to this subspecies, which was previously united with anatum. It is the New World equivalent to calidus. It is smaller than anatum. It is also paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus.[29] Juveniles are browner, and less grey, than in calidus, and paler, sometimes almost sandy, than in anatum.
- Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary Falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands, and is non-migratory;[12] it is endangered with only six to eight pairs surviving.[5] Males have a rufous wash on crown, nape, ears and back; underside conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.[12]
- Falco peregrinus minor was first described by Bonaparte in 1850. It was formerly often perconfusus.[30] It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory, and small and dark.
- Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and Comoros. It is non-migratory.[12]
- Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean Peregrine Falcon or the Maltese Falcon.[31] It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides, Barbary Falcons (see also below), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies, and the underside usually has rusty hue.[12] Males weigh around 445 g, while females weigh up to 920 g.[6]
- Falco peregrinus ernesti, described by Sharpe in 1894, is found from Indonesia to Philippines and south to Papua New Guinea and Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate in the very dark, dense barring on its underside and its black ear coverts.
- Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands. It is non-migratory. It is very rare, and may only remain on a single island.[5] It is a dark form, resembling pealei in color, but darker, especially on tail.[12]
- Falco peregrinus nesiotes described by Mayr in 1941,[32] is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.[33]
- Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through north Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g.[6]
- Falco peregrinus babylonicus described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and Tian Shan to Mongolian Altai ranges. It is paler than pelegrinoides, and somewhat similar to a small, pale Lanner Falcon. It is smaller than Peregrine Falcon; males weigh 330–400 g, while females weigh 513–765 g.[6]
These last two races are often split as Barbary Falcon Falco pelegrinoides.[6] There is a 0.6–0.7% genetic distance in the Peregine-Barbary Falcon ("peregrinoid") complex.[34] These birds inhabit arid regions from the Canary Islands along the rim of the Sahara through the Middle East to Central Asia and Mongolia. They have a red neck patch but otherwise differ in appearance from the Peregrine proper merely according to Gloger's Rule.[35] The Barbary Falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the Peregrine, but less often and far less pronounced.[6] The Barbary Falcon's shoulder and pelvis bones are stout by comparison with the Peregrine, and its feet are smaller.[36] They have no postzygotic reproduction barriers in place,[37] but they breed at different times of year than neighboring Peregrine Falcon subspecies.[21][38][6][39][40][34][41]
[ Ecology and behavior
The Peregrine Falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities.[12] In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climes typically migrate great distances during the northern winter.[42] The Peregrine Falcon is often stated to be the fastest animal on the planet in its hunting dive, the stoop, which involves soaring to a great height and then diving steeply at speeds commonly said to be over 320 km/h (199 mph), and hitting one wing of its prey so as not to harm itself on impact.[4] A study testing the flight physics of an 'ideal falcon' found a theoretical speed limit at 400 km/h (250 mph) for low altitude flight and 625 km/h (390 mph) for high altitude flight.[43] In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).[44]
The life span in the wild is up to 15.5 years.[6] Mortality in the first year is between 59–70%, declining to between 25–32% in adults.[6] Apart from anthropogenic threats like collision with human-made objects, the Peregrine may be killed by large eagles or large owls.[25] The Peregrine Falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the Peregrine Falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known Peregrine Falcon ectoparasites are chewing lice[45] Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).[46]
[ Feeding
The Peregrine Falcon feeds almost exclusively on medium sized birds such as doves, waterfowl, songbirds, waders and pigeons.[14] Worldwide, it is estimated that between 1,500 and 2,000 bird species are predated by these falcons. In North America, prey has varied in size from 3-g hummingbirds to a 3.1-kg Sandhill Crane (killed by a peregrine in a swoop).[47] Other than bats taken at night,[48] it rarely hunts small mammals, but will on occasion take rats, voles, hares, mice and squirrels; the coastal populations of the large subspecies pealei feed almost exclusively on seabirds.[13] In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed while successfully hunting a juvenile Scarlet Ibis.[49] Insects and reptiles make up a small proportion of the diet, which varies greatly depending on what prey is available.[14] In urban areas, the main item of the Peregrine's diet is the Rock or Feral Pigeon, which comprise 80% or more of the dietary intake for peregrines in some cities. Other common city birds are also taken regularly, such as Mourning Doves, Common Swifts, Northern Flickers, Common Starlings, American Robins and various corvids.[48]
The Peregrine Falcon hunts at dawn and dusk, when prey are most active, but in cities also nocturnally, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by Peregrines include species as diverse as Yellow-billed Cuckoo, Black-necked Grebe, Virginia Rail and Common Quail.[48] It requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields and tundra. It searches for prey either from a high perch or from the air.[50] Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked.[13] The air pressure from a 200 mph (320 km/h) dive could possibly damage a bird's lungs, but small bony tubercles in a falcon's nostrils guide the shock waves of the air entering the nostrils (compare intake ramps and inlet cones of jet engines), enabling the bird to breathe more easily while diving by reducing the change in air pressure.[51] To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. Prey is struck and captured in mid-air; the Peregrine Falcon strikes its prey with a clenched foot, stunning or killing it, then turns to catch it in mid-air.[50] The Peregrine will drop it to the ground and eat it there if it is too heavy to carry. Prey is plucked before consumption.[51]
[ Reproduction
The Peregrine Falcon is sexually mature at the end of the first year of age but in healthy populations they breed after two to three years of age. The pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives.[10] The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons. The Peregrine Falcon is territorial during the breeding season; nesting pairs are usually more than 1 km (0.6 miles) apart, and often much farther, even in areas with large numbers of pairs.[52] The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two to seven in a 16 year period. The pair defends the chosen nest site against other Peregrines, and often against ravens, herons, gulls and (in ground nest) mammals like foxes, wolverines, felids, bears and wolves.[52] Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or Gyrfalcons. Peregrines defending their nests have managed to kill raptors as large as Golden Eagles and Bald Eagles (which they normally avoid) that have come close to the nest.[53]
The Peregrine Falcon nests in a scrape, normally on cliff edges or, today regularly in many parts of its range, on tall buildings or bridges. Cliff nests are generally located under an overhang, on ledges with vegetation, and south-facing sites are favored.[13] In some regions, as in parts of Australia and on the west coast of Northern North-America, large tree hollows are used for nesting. Before the demise of most European peregrines, there was a large population of peregrines in central and western Europe using the disused nests of other large birds.[14] The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added.[10] In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. The human-made structures used for breeding closely resemble the natural cliff ledges that the Peregrine prefers for its nesting locations.[5][52]
Mostly three to four eggs (range 1-5) are laid in the scrape.[54] The eggs are white to buff with red or brown markings.[54] They are incubated for 29 to 33 days, mainly by the female.[13] The male also helps with the incubation of the eggs over day, but at night only the female incubates. The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere (thee Australian subspecies macropus may breed as late as November and equatorial populations may nest anytime between June and December). The female generally lays a
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